Cirsium occidentale var. coulteri

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Variety coulteri grows in the coastal zone of central and southern California. The epithet coulteri was for many years misapplied by California botanists. W. L. Jepson ([1923–1925]) used it at the varietal level for a range of plants that included parts of what I am calling vars. venustum and californicum. P. A. Munz (1959) and various other authors recognized Cirsium coulteri as a species, primarily in the context that I recognize as var. venustum. J. T. Howell (1959b) examined the type of C. coulteri and concluded that it and C. occidentale (in the strict sense) are synonymous. I have examined photos of the types of both C. occidentale and C. coulteri and have reached a different conclusion. The type of C. occidentale (BM) has heads with ± imbricate, comparatively short phyllary appendages. The type of C. coulteri (TCD) has long, acicular, subequal phyllaries. D. J. Keil and C. E. Turner (1993) treated the plants with the two head types all as var. occidentale, but I believe it is preferable to recognize separate varieties. Some intermediates are known, but var. coulteri is usually readily separable from var. occidentale.